There are 5 skulls that can be found in the dungeons.

Location Walkthrough
Level 2

Secret 4

After passing through the Teleport after the Crystal of Life, and then two more doors, there is a lever in a room to the left. Pulling it will open another room on the left, before the one with the lever. The skull is inside a Phalanx Helmet in the back corner.
Level 3 After the first puzzle is an area with some pits. The skull is on the floor in the furthest corner of the room at the bottom of the pits (technically therefore being on level 4), behind where the Teleport appears upon pulling the lever.
Level 7 The skull is in an alcove near the normal stairs down to level 8.
Level 8 This skull is found on the floor in a room to which you must teleport (from the southeastern tunnels of the map) where you must fight a number of Herders in order to find a key.
Level 10 This skull is found in an area beneath the pits in the northern part of the level (and is therefore technically found on level 11).

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